Date of Award

Summer 7-21-2025

Embargo Period

8-6-2027

Document Type

Dissertation - MUSC Only

Degree Name

Doctor of Philosophy (PhD)

Department

Molecular and Cellular Biology and Pathology

College

College of Graduate Studies

First Advisor

Adviye Ergul

Second Advisor

Onder Albayram

Abstract

Palmitic acid (PA) is the most abundant saturated fatty acid in the human body, playing a critical role in multiple metabolic processes. Epidemiological studies unequivocally demonstrate that diets high in saturated fatty acids and elevated levels of palmitic acid are directly associated with diminished cognitive function. PA is sourced both from dietary intake and synthesized through de novo lipogenesis in the liver, and it possesses the remarkable ability to cross the blood-brain barrier. This capability increases the fatty acid pool within the brain, ultimately influencing cognitive health. Importantly, palmitic acid (C16:0) has been linked to severe neurodegenerative processes, most notably to the neuronal hyperphosphorylation and deposition of tau protein. Given that cerebrovascular dysfunction is one of the earliest pathologies detected in dementia, our novel hypothesis is that a PA-rich diet mediates neurovascular dysfunction in part by cerebrovascular tau hyperphosphorylation without neurodegeneration, leading to cognitive impairment. To rigorously test this hypothesis, we implemented a comprehensive approach to evaluate both physiological and cognitive functions over time, supported by lipidomic, transcriptomic, molecular, and biochemical studies. In a preclinical murine model exposed to a novel PA-rich diet, significant cerebrovascular tau phosphorylation (p-tau), neurovascular uncoupling, and behavioral dysfunction were observed as early as twelve weeks. Over time, the behavioral dysfunction progressed into cognitive impairment, coupled with sustained structural and functional cerebrovascular deficits as determined by transmission electron microscopy and Laser Doppler flowmetry. PA-rich diet didn’t promote cerebrovascular p-tau accumulation associated behavioral deficits in a global tau knockout (KO) murine model, underscoring the importance of p-tau accumulation in the cerebrovasculature. Moreover, another transgenic animal harboring the humanized tau has shown accelerated cognitive and neurovascular impairment after being on the same PA-rich diet for 12 weeks. Collectively, using a PA-rich diet in three different mouse models and integrating molecular, biochemical, and functional methodologies, we provide novel evidence that dysregulation of tau protein in the cerebrovasculature leads to structural and functional impairments, which are likely precursors to neurodegeneration.

Rights

Copyright is held by the author. All rights reserved.

Available for download on Friday, August 06, 2027

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